Lein graduated from UCLA with undergraduate and doctorate degrees in zoology and a focus on endocrinology. He taught at the Ohio State University and Vanderbilt medical schools before becoming an assistant professor in the physiology department of the Northwestern University Medical School in 1947. There, he later became professor and director of student affairs, and finally was named assistant dean. During this time he spent a year as a Guggenheim Fellow at the Laboratoire de Biochimie, College de France.

Lein was respected for his research in endocrinology, and in 1979 published a book titled ""The Cycling Female: Her Menstrual Rhythm."" The book was praised by reviewers for being ""written in a warm and comfortable style without sacrificing biological accuracy or detail.""

Lein is survived by his wife Teresa; his daughter Laura Lein of Austin, Tex.; his son David Lein of Berkeley, Calif.; three grandchildren, Anna, Rebecca and David Kuipers; and nephew, Benjamin Lein, of Los Angeles.

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UC San Diego's independent student newspaper since 1967

The UCSD Guardian

UC San Diego's independent student newspaper since 1967

The UCSD Guardian

UC San Diego's independent student newspaper since 1967

The UCSD Guardian